《中国植物志》|| 忍冬科 CAPRIFOLIACEAE ||合轴组 Sect. 2. Pseudotinus C. B. Clarke ||显脉荚蒾 Viburnum nervosum D. Don
13. 显脉荚蒾（拉汉种子植物名称）心叶荚蒾（拉汉种子植物名称）图版7：1-4
Viburnum nervosum D. Don, Prodr. Fl. Nepal. 141. 1825; Stapf in Bot. Mag. 150: t. 9063, in textu. 1925; Hara in Journ. Jap. Bot. 47: 275, 1972, et Fl. East. Himal. 3: 107. 1975. ——V. cordifolium Wall. ［Cat. no. 462. 1829, nom. nud. ; ex DC. Prodr. 4: 327. 1830, descr. ; Hook. f. et Thoms. in Journ. Linn. Soc. 2: 175. 1858, excl. syn. ; 中国高等植物图鉴4: 313. 1975. ——V. furcatum auct. non Blume: Hook. f. et Thoms. in l. c., pro syn. sub V. cordifolium Wall. ——V. sympodiale auct. non Graebn. : Hand. -Mazz. Symb. Sin. 7: 1035. 1936, pro parte.
落叶灌木或小乔木，高达5米；幼枝、叶下面中脉和侧脉上、叶柄和花序均疏被鳞片状或糠粃状簇状毛；二年生小枝灰色或灰褐色，无毛，具少数大形皮孔。叶纸质，卵形至宽卵形，稀矩圆状卵形，长9-18厘米，顶端渐尖，基部心形或圆形，边缘常有不整齐钝或圆的锯齿，很少为尖锯齿，上面无毛或近无毛，下面常多少被簇状毛，侧脉8-10对，上面凹陷，下面凸起，小脉横列；叶柄粗壮，长2-5.5厘米，有或无托叶。聚伞花序与叶同时开放，直径5-15厘米，无大型的不孕花，连同萼筒均有红褐色小腺体，第一级辐射枝5-7条，花生于第二至第三级辐射枝上；萼筒筒状钟形，长约1.5毫米，无毛，萼齿卵形，被少数簇状毛；花冠白色或带微红，辐状，直径6-10毫米，裂片长为筒的2倍，卵状矩圆形至矩圆形，大小常不等，外侧者常较大，尤以花序边缘的花为甚；雄蕊花丝长约1毫米，花药宽卵圆形，紫色；花柱略高出萼齿。果实先红色后变黑色，卵圆形，长约8毫米，直径6-7毫米；核扁，两缘内弯，有1条浅背沟和1条深腹沟。花期4-6月；果熟期9-10月。
产湖南南部（天堂山），广西东北部（临桂），四川西部和西南部，云南西北部（南达景东）、西部和东北部及西藏南部至东南部。生于山顶或山坡林中和林缘灌丛中，冷杉林下常见，海拔 (1800-) 2100-4500米。印度、尼泊尔、锡金、不丹、缅甸北部和越南北部也有分布。
Viburnum cordifolium Wall. ex DC. var. hypsophilum Hand. -Mazz. Symb. Sin. 7: 1034. 1936的模式标本 H. Handel-Mazzetti 9293，系采自云南西北部。其主要特征为：植物体较小，高不到1米，叶较小，长3-6厘米，顶圆或有时微凸头，花序小，仅有1-8朵花。我们怀疑这个变种可能是由环境因素所造成的可塑性变化。姑志于此，留待考订。
Source 中国植物物种信息数据库
《云南植物志》|| 忍冬科 Caprifoliaceae nom. conserv. ||荚蒾属 Viburnum L. ||心叶荚蒾 Viburnum nervosum D. Don
4．心叶荚蒾（图鉴）
维塞夫（纳西族语译音）
Viburnum nervosum D. Don (1825); Stapf in Curtis (1925); Hara (1972), (1975), (1979).
Viburnum cordifolium Wall. ex A. DC. (1830); Hook. f. et Thoms. (1858); Franch. (1885); Rehd. in Sarg. (1908)*, (1911), (1940); Hand.-Mazz. (1936); Hara (1966), (1971); 图鉴 (1975), in nota; ? V. sympodiale auct. non Graebn.: Hand.-Mazz. (1936), p. p., quoad specim. Yunnan.
落叶灌木至小乔木，高1—5米；幼枝、叶片背面脉上、叶柄及花序均被黄褐色簇毛状鳞片；二年生小枝平滑而棕褐色，疏生皮孔。冬芽无鳞片。叶片近坚纸质，轮廓椭圆形，长（4—）7—15厘米，宽（2.5—）4—11厘米，先端短渐尖或急尖，基部心形或有时圆形，边缘有不整齐的锐锯齿，叶面深绿色，背面淡绿色，侧脉每边6—8条，伸达齿端；叶柄长1.5—3厘米，腹面具槽，无托叶。花序聚伞状复伞形，无总梗，长2—5（6）厘米，宽3.5—5（—10）厘米，第一级辐射枝5—6（—7）条；苞片和小苞片线状披针形，长5—6（—8）毫米，外被黄褐色簇毛状鳞片，早落。花芳香，着生于第二级辐射枝上，无梗；萼筒长约2.5毫米，无毛，萼檐5齿，齿披针形，有簇状毛；花冠白色，辐状，直径达8毫米，冠筒极短，花冠裂片长4毫米；雄蕊5，长不及花冠之半。核果椭圆形，长达8毫米，红色而后变黑色；核略扁，背具1浅槽，腹具1深沟。花期4—6月，果期7—9月。
产滇西北和东北，而南达景东，滇东南的文山，生于海拔（1500—）2450—4000米之山谷及山坡林内或灌丛中，冷杉林下尤其常见；分布于湖南（天堂山）、广西（临桂）、四川西部、以及西藏东南部。印度东北部、锡金、不丹、缅甸北部及越南北部也有。
Source 中国植物物种信息数据库
《Flora of China》|| Adoxaceae || Viburnum ||显脉荚蒾 Viburnum nervosum
14. Viburnum nervosum D. Don, Prodr. Fl. Nepal. 141. 1825.
显脉荚蒾 xian mai jia mi
Solenotinus nervosus (D. Don) Oersted; Viburnum cordifolium Wallich ex Candolle; V. cordifolium var. hypsophilum Handel-Mazzetti; V. nervosum var. hypsophilum (Handel-Mazzetti) H. W. Li.
Shrubs or small trees, deciduous, to 5 m tall. Bark gray-brownish. Branchlets of current year lepidote or furfuraceous stellate-pubescent; branchlets of previous year gray or gray-brownish, terete, glabrous, with sparse, dispersed, large, elliptic lenticels. Winter buds naked, furfuraceous stellate-pubescent. Leaves often clustered at apices of branchlets; stipules 2, subulate, 2-5 mm, caducous, or absent; petiole green, robust, 2-5.5 cm, brownish stellate-pubescent; leaf blade greenish brown when young, ovate to broadly ovate, rarely oblong-ovate, (4-)7-18 × (2.5-)4-11 cm, papery, abaxially often ± stellate-pubescent particularly along veins, adaxially glabrous or subglabrous, midvein raised abaxially, lateral veins 8-10-jugate, pinnate, arched, branched, anastomosing near margin, raised abaxially, impressed adaxially, veinlets transverse, conspicuous on both surfaces, not lobed, base cordate or rounded, without glands, margin irregularly serrate, apex acuminate. Flowers appearing with leaves; inflorescence a compound umbel-like cyme, terminal, 5-15 cm in diam.; rays whorled; first node of inflorescence with 5-7 rays, dense, with small red-brownish glands, without large sterile radiant flowers; peduncles absent; bracts caducous, leaflike, green, ovate to elliptic, stellate-pubescent; bracteoles scalelike. Flowers on rays of 2nd and 3rd orders, fragrant, subsessile or shortly pedicellate. Calyx greenish; tube tubular-campanulate, ca. 1.5 mm, with small red-brownish glands; lobes ovate, very small, 0.5-1 mm, sparsely stellate-pubescent, apex obtuse. Corolla white or reddish, rotate, 5-8 mm in diam., glabrous; tube 3-4 mm; lobes spreading, ovate-oblong to oblong, ca. 2 × as long as tube, unequal in size, outer ones, particularly marginal ones, often larger, apex rounded, margin entire. Stamens shorter than corolla, inserted near base of corolla tube; filaments ca. 1 mm; anthers purple, broadly ovoid, ca. 1 mm. Styles slightly exceeding calyx lobes; stigmas capitate. Fruit initially turning yellow, then red, maturing purplish black, ovoid, 7-9 × 5-7 mm, base rounded, apex rounded, glabrous; pyrenes compressed, oblong, 6-8 × 4-5 mm, with 1 shallow dorsal groove and 1 deep ventral groove, apex rounded. Fl. Apr-Jun, fr. Sep-Oct. 2n = 18.
Forests (usually Abies forests), scrub; (1800-)2100-4500 m. Sichuan, Xizang, Yunnan [Bhutan, India, N Myanmar, Nepal, N Vietnam].
Source eFloras.org
