《中国植物志》|| 茜草科 RUBIACEAE ||玉叶金花属 Mussaenda Linn. ||玉叶金花 Mussaenda pubescens W. T. Aiton

17. 玉叶金花（广东）野白纸扇（广州）、良口茶（广东）
Mussaenda pubescens Ait. f. Hort. Kew. ed. 2, 1: 372. 1810; DC. Prodr. 4: 371. 1830; Benth. Fl. Hongk. 153. 1861; Hemsl. in Journ. Linn. Boc. Bot. 23: 379. 1888; Matsum. et Hayata, Enum. Pl. Formosa. 188. 1906; Hayata, Ic. Pl. Formos. 2: 94. 1912; Rehd. in Journ. Arn. Arb. 16: 320. 1935; Kanehira, Formos. Trees ed. 2, 674, t. 628. 1936; Masam. in Trans. Nat. Hist. Soc. Formosa. 28: 117. 1938; 广州植物志 506. 1956; Li, Woody Fl. Taiwan 863, fig. 348. 1963; 海南植物志3: 314. 1974; Ferguson in Not. Roy. Bot. Gard. Edinb. 32 (1): 110. 1972; 中国高等植物图鉴4: 195, 图5804. 1975; 云南种子植物名录, 下册: 1263. 1984.——M. bodinieri Levl. in Bull. Soc. Bot. France 55: 59. 1908; Hutchins. in Sargent, Pl. Wils. 3: 396. 1916.
攀援灌木，嫩枝被贴伏短柔毛。叶对生或轮生，膜质或薄纸质，卵状长圆形或卵状披针形，长5-8厘米，宽2-2.5厘米，顶端渐尖，基部楔形，上面近无毛或疏被毛，下面密被短柔毛；叶柄长3-8毫米，被柔毛；托叶三角形，长5-7毫米，深2裂，裂片钻形，长4-6毫米。聚伞花序顶生，密花；苞片线形，有硬毛，长约4毫米；花梗极短或无梗；花萼管陀螺形，长3-4毫米，被柔毛，萼裂片线形，通常比花萼管长2倍以上，基部密被柔毛，向上毛渐稀疏；花叶阔椭圆形，长2.5-5厘米，宽2-3.5厘米，有纵脉5-7条，顶端钝或短尖，基部狭窄，柄长1-2.8厘米，两面被柔毛；花冠黄色，花冠管长约2厘米，外面被贴伏短柔毛，内面喉部密被棒形毛，花冠裂片长圆状披针形，长约4毫米，渐尖，内面密生金黄色小疣突；花柱短，内藏。浆果近球形，长8-10毫米，直径6-7.5毫米，疏被柔毛，顶部有萼檐脱落后的环状疤痕，干时黑色，果柄长4-5毫米，疏被毛。花期6-7月。
产于广东、香港、海南、广西、福建、湖南、江西、浙江和台湾。生于灌丛、溪谷、山坡或村旁。模式标本采自我国南部，具体地点不详。
茎叶味甘、性凉，有清凉消暑、清热疏风的功效，供药用或晒干代茶叶饮用。

《云南植物志》|| 茜草科 Rubiaceae ||玉叶金花属 Mussaenda Linn. ||玉叶金花 Mussaenda pubescens Ait. f.

13.玉叶金花（广东）
白纸扇（云南种子植物名录）
Mussaenda pubescens Ait. f. (1810); Hutch. in Sargent (1916); Ferguson in Lauener (1972); 图鉴 (1975)*; 云南种子植物名录 (1984); 中国植物志 (1999).
Mussaenda bodinieri Lévl. (1908), (1917); Hutch. in Sargent (1916).
攀援灌木。嫩枝被贴伏短柔毛。叶对生或轮生，膜质或薄纸质，卵状长圆形或卵状披针形，长5—8厘米，宽2—2.5厘米，先端渐尖，基部楔形，叶面近无毛或疏被柔毛，背面密被短柔毛；叶柄长3—8毫米，被柔毛；托叶三角形，长5—7毫米，2深裂，裂片钻形，长4—6毫米。聚伞花序顶生，密花；苞片线形，有硬毛，长约4毫米；花梗极短或无梗；萼管陀螺形，长3—4毫米，被柔毛，萼裂片线形，常比萼管长2倍以上，基部密被柔毛，向上毛渐稀疏；花叶阔椭圆形，长2.5—5厘米，宽2—3.5厘米，有纵脉5—7条，先端钝或短尖，基部渐狭，柄长1—2.8厘米，两面被柔毛；花冠黄色，冠管长约2厘米，外面被贴伏短柔毛，冠管喉部密被棒状毛，花冠裂片长圆状披针形，长约4毫米，内面有密的金黄色小疣突；花柱短，内藏。浆果近球形，长8—10毫米，直径6—7.5毫米，疏被柔毛，顶部有环状疤痕，干时黑色，果柄长4—5毫米，疏被柔毛。 花期6—7月。
产绥江、大关、师宗、新平、元江、峨山、文山、金平、绿春、思茅、勐腊、景洪、勐海、盈江、潞西、瑞丽；生于海拔1200—1500米处的沟谷或旷野灌丛。分布于贵州、广西、广东、香港、海南、湖南、江西、福建、浙江、台湾。
茎叶及根均可药用，味甘、性凉，有清凉消暑、清热疏风的功效，可治感冒、中暑、支气管炎、水肿；叶晒干可做茶叶的代用品。

《Flora of China》|| Rubiaceae || Mussaenda ||玉叶金花 Mussaenda pubescens

25. Mussaenda pubescens W. T. Aiton, Hort. Kew., ed. 2. 1: 372. 1810.

玉叶金花 yu ye jin hua

Mussaenda bodinieri H. Léveillé & Vaniot; M. pubescens var. alba X. F. Deng & D. X. Zhang; M. pubescens f. clematidiflora Chun ex H. H. Hsue & H. Wu.

Climbing shrubs, often extensively twining; branches terete, densely strigillose and sometimes also villosulous, sometimes with congested groups of leaves borne on reduced axillary buds. Leaves opposite or perhaps rarely whorled; petiole 3-8 mm, moderately to densely strigillose; blade drying membranous or thinly papery, ovate-oblong, ovate-lanceolate, elliptic, lanceolate, or oblanceolate, 2-9 × 1-4 cm, adaxially sparsely strigillose to glabrescent on lamina and moderately to densely strigillose on veins, abaxially sparsely to densely strigillose with pubescence denser on veins, base acute to obtuse, apex acute to slightly acuminate; secondary veins 4-7 pairs, tertiary venation reticulate; stipules usually deciduous, triangular, 3-7 mm, moderately to densely strigillose, deeply 2-lobed, lobes narrowly triangular to subulate. Inflorescences terminal, subcapitate to congested-cymose, unbranched or sometimes tripartite, 1-3 × 1-3 cm, densely strigillose to villosulous, sessile to pedunculate; peduncles 0.1-1.4 cm; bracts linear, 3-5 mm; pedicels to 1 mm. Flowers sessile or infrequently pedicellate, biology not noted. Calyx moderately to densely strigillose to strigose; hypanthium portion ellipsoid to turbinate, 1.5-3 mm; lobes linear to narrowly triangular, 3-6 mm, with 1(or 5) lobes on 1-3 flowers per inflorescence of some plants expanded into white calycophyll, blade elliptic, broadly elliptic, ovate, or lanceolate, (0.4-)2.5-5 × (0.2-)2-3.5 cm, sparsely to moderately puberulent or strigillose on both surfaces, base acute to rounded, stipe 3-28 mm, apex obtuse to acute. Corolla white or yellow, salverform, outside moderately to densely strigillose or strigose; tube 11-20 mm, densely clavate pubescent in throat; lobes oblong-lanceolate to lanceolate, 2.5-4 mm, adaxially densely golden yellow papillose, acuminate. Berry subglobose, 8-10 × 6-7.5 mm, sparsely strigillose to glabrescent, smooth or finely lenticellate, calyx limb caducous, sometimes stipitate with stipe to 5 mm. Fl. Apr-Jul, fr. Jun-Dec.

Thickets in ravines, on hill slopes, or at village margins or roadsides; below 100-900 m. Fujian, Guangdong, Guangxi, Hainan, Hunan, Jiangxi, Taiwan, Zhejiang [Vietnam].

This species is widespread, common, and morphologically variable. H. H. Hsue and H. Wu (in FRPS 71(1): 296. 1999) described the leaf arrangement as opposite or whorled, but only opposite leaves have been seen on the numerous specimens studied.

One plant from Guangdong (Gaoyao) has all five of the calyx lobes petaloid and enlarged on each flower, though these structures are smaller than calycophylls that are borne singly on a flower. These plants have been separated as Mussaenda pubescens f. clematidiflora; this case was studied by Deng and Zhang (Acta Phytotax. Sin. 44: 611. 2006), who concluded that this plant is better regarded as developmentally abnormal and formally synonymized this name here. Plants with white corollas have been separated as M. pubescens var. alba; at least a third of the specimens of M. pubescens studied belong to this group, and considering that this color variation is common in other Rubiaceae species the variety is not recognized taxonomically here.